In the tessellation there are two families of mudskippers, aquatic and terrestrial. As the tide advances and recedes, the two worlds intersect through the rippled mud. On sea and on the land, a pair square off in a display of aggression.

Richard notes: In August 2015, a new pentagonal tiling made global news, including the BBC and the Guardian website. I found this very exciting, and decided to test out if I could make a tessellation on it, treating it as a speed challenge. I developed this tessellation within a few days, and completed the coloured version within the month. The tile had an interesting kind of rippling structure, this reminded me of ripples in the mangroves, and I have always had a soft spot for the Goby family of fish, with their comically sad faces and bulging eyes, small limbs and often spectacular dorsal fins and colouring, and their aggressive display rituals. I wish that they had been the original creatures to emerge from the sea, imagine what bizarre forms terrestrial life may have evolved to based on this template!


There are two tile designs. While the pentagonal tiling as published is a single tile with a  mirror image, once the edge profiles are solved, the tessellation tiles are not mirror images. The overall tiling is not aperiodic, but it is anisohedral, each tile appears in several orientations and relationships to its surrounding tiles.

Mudskipping Tiles

Tiles for the tessellation Mudskipping

Mudskipping detail

Mudskipping – detail of the print